Orchids are extraordinary plants that have evolved the strategy of dispersing their pollen in little sacs called pollinia. Pollinia are normally attached by supports (caudicles) to adhesive pads (viscidia) that stick to various body parts of the pollinator. The entire pollination unit is called a pollinarium. Since pollen dispensed in pollinia is not available as food to most insects ( van der Cingel 1995 ), orchids supply nectar or have evolved ingenious patterns of mimicry and deceit to attract potential pollinators ( Endress 1994 ).

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Beetle associations with orchids are diverse. Beetles may visit orchid flowers simply for nectar, or at the same time acquire pollinaria and serve as pollinators. Beetles can also feed and raise their brood on orchid floral and vegetative parts. Some orchids depend on various Coleoptera for pollination and have produced “cantharophilous” flowers especially attractive to beetles ( Endress 1994...

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